Peer-reviewed journal articles
You will find links to all PDFs as hosted at the journals' websites. I can also send PDFs personally to colleagues by e-mail request - bruno.buzatto [at]

2014 - present (post-PhD)

41. Lymbery SJ, Tomkins JL, Buzatto BA & DJ Hosken (2021). Kin-mediated plasticity in alternative reproductive tactics. Proc R Soc B-Biol Sci, In press.

40. Machado G, Buzatto BA & DSM Samia (2021). It is not always about body size: evidence of Rensch’s rule in a male weapon indoors? Biology Letters, 17(6): 20200767. [PDF at Biology Letters]

39. Buzatto BA & L Haeusler & N Tamang (2021). Trapped indoors? Long-distance dispersal in Mygalomorph spiders and its effect on species ranges. Journal of Comparative Physiology A, 207: 279-292. [PDF]

38. Buzatto BA & HL Clark (2020) Selection for male weapons boosts female fecundity, eliminating sexual conflict. Scientific Reports, 10(2311): 1-7. [PDF at Sci Reports]

37. Firman RC, Rubenstein DR, Moran JM, Rowe KC & BA Buzatto (2020) Extreme and variable climatic conditions drive the evolution of sociality in Australian rodents. Current Biology, 30(4): 691-697.e3  [PDF at Current Biology]

36. Buzatto BA, JS Kotiaho, LAF Assis & LW Simmons (2019) A link between heritable parasite resistance and mate choice in dung beetles. Behavioral Ecology, 30(5): 1382-1387. [PDF at Behav Ecol]

35. Eberhard WG, RL Rodríguez, BA Huber, B Speck, H Miller, BA Buzatto & G Machado (2018) Sexual selection and static allometry: the importance of function. The Quarterly Review of Biology, 93(3): 207-250. [PDF at Q Rev Biol]

34. Buzatto BA, HL Clark & JL Tomkins (2018) Morph-specific artificial selection reveals a constraint on the evolution of polyphenisms. Proc R Soc B-Biol Sci, 285(1879): 20180335 [PDF at Proc R Soc B]

33. McCullough EL, BA Buzatto & LW Simmons (2018) Population density mediates the interaction between pre- and post-mating sexual selection. Evolution, 72(4): 893-905. [PDF at Evolution].

32. Pike KN, JL Tomkins & BA Buzatto (2017) Mixed evidence for the erosion of inter-tactical genetic correlations through intralocus tactical conflict. J Evol Biol, 30(6): 1195-1204. [PDF at J Evol Biol]

31. McCullough EL,
BA Buzatto & LW Simmons (2017) Benefits of polyandry: molecular evidence from field-caught dung beetles. Molecular Ecology, 26(13): 3546-3555.
[PDF at Mol Ecol]

30. Rowland JM, CR Qualls &
BA Buzatto (2017) A model for conditional male trimorphisms. J Theor Biol, 419: 184-192. [PDF at J Theor Biol]

29. Buzatto BA, EM Thyer, JD Roberts & LW Simmons (2017) Sperm competition and the evolution of precopulatory weapons: testis size and amplexus position, but not arm strength, affect fertilization success in a chorusing frog. Evolution
, 71(2): 329-341. [PDF at Evolution]

28. Machado G, BA Buzatto, S García-Hernández & R Macías-Ordóñez (2016) Macroecology of sexual selection: a predictive conceptual framework for large-scale variation in reproductive traits. Am Nat, 188: S8–S27. [PDF at Am Nat]

27. Silva DP, B Vilela, BA Buzatto, AP Moczek & J Hortal (2016) Contextualized niche shifts upon independent invasions by the dung beetle Onthophagus taurus. Biol Invasions, 18(11):3137-3148. [PDF]

26. Buzatto BA, M Buoro, WN Hazel & JL Tomkins (2015) Investigating the genetic architecture of conditional strategies using the Environmental Threshold Model. Proc R Soc B-Biol Sci, 282(1821): 20152075 [PDF at Proc R Soc B]

25. Buzatto BA, JD Roberts & LW Simmons (2015) Sperm competition and the evolution of pre-copulatory weapons: increasing male density promotes sperm competition and reduces selection on arm strength in a chorusing frog. Evolution, 69(10): 2613-2624. [PDF at Evolution]
Featured in the ABC and Science Network Western Australia  [links to the stories here]

24. Buzatto BA, JS Kotiaho, JL Tomkins & LW Simmons (2015) Intralocus tactical conflict: genetic correlations between fighters and sneakers of the dung beetle Onthophagus taurusJ Evol Biol, 28(3): 730-738. 
[PDF at J Evol Biol]

23. Muniz DG, PR Guimarães, BA Buzatto & G Machado (2015) A sexual network approach to sperm competition in a species with alternative mating tactics. Behav Ecol, 26(1): 121-129. [PDF at Behavioral Ecology]

22. Buzatto BA & G Machado (2014) Male dimorphism and alternative reproductive tactics in harvestmen (Arachnida: Opiliones). Behav Process, 109 (Part A): 2-13. [PDF at Behavioural Processes]

21. Wilson CJ, BA Buzatto, SP Robinson & JL Tomkins (2014) Sociosexual environment influences patterns of ejaculate transfer and female kicking in Callosobruchus maculatus. Anim Behav, 94: 37-43.
[PDF at Animal Behaviour]

20. Buzatto BA, WN Hazel & JL Tomkins (2014) A theoretical muddle of the conditional strategy: a comment on Neff and Svensson. Philos Trans R Soc B-Biol Sci, 369(1642): SI; Article Number: 20130625.
[PDF at Phil Trans R Soc B]
First indexed comment ever published by this journal

19. Buzatto BA, JL Tomkins, LW Simmons & G Machado (2014) Correlated evolution of sexual dimorphism and male dimorphism in a clade of Neotropical harvestmen. Evolution, 68(6): 1671-1686.
[PDF at Evolution]

18. Simmons LW &
BA Buzatto (2014). Contrasting responses of pre- and post-copulatory traits to variation in mating competition. Funct Ecol, 28(2): 494–499.
[PDF at Functional Ecology]

2009 - 2013 (during PhD)

17. Santos AJ, RL Ferreira &
BA Buzatto (2013) Two new cave-dwelling species of the short-tailed whipscorpion genus Rowlandius (Arachnida: Schizomida: Hubbardiidae) from northeastern Brazil, with comments on male dimorphism. PLoS One, 8(5): e63616.
[PDF] [at PLoS One]
Featured in the Brazilian newspaper Estado de Minas  [at Estado de Minas]

16. Requena GS, BA Buzatto, EG Martins & G Machado (2012) Paternal care decreases foraging activity and body condition, but does not impose survival costs to caring males in a Neotropical arachnid. PLoS One, 7(10): e46701. [PDF] [at PLoS One]
Featured in The New York Times  [at The New York Times]

15. Buzatto BA, JL Tomkins & LW Simmons (2012) Maternal effects on male weaponry: female dung beetles produce major sons with longer horns when they perceive higher population density. BMC Evol Biol, 12: 118. [PDF] [at BMC Evol Biol]           Featured in the magazine The Scientist  [story at The Scientist] [interview with Russell Woolf, ABC Radio Perth]

14. R Munguía-Steyer, BA Buzatto & G Machado (2012) Male dimorphism in a Neotropical arachnid: harem size, sneaker opportunities and gonadal investment. Behav Ecol, 23(4): 827–835. [PDF at Behavioral Ecology]

13. Buzatto BA, LW Simmons & JL Tomkins (2012) Paternal effects on the expression of a male polyphenism. Evolution, 66(10): 3167–3178.
[PDF at Evolution]

12. Buzatto BA, LW Simmons & JL Tomkins (2012) Genetic variation underlying the expression of a polyphenism. J Evol Biol, 25: 748–758.
[PDF at J Evol Biol]

11. Buzatto BA, GS Requena, RS Lourenço, R Munguía-Steyer & G Machado (2011) Conditional male dimorphism and alternative reproductive tactics in a Neotropical arachnid (Opiliones). Evol Ecol, 25: 331– 349. [PDF at Evol Ecol]

10. Requena GS, BA Buzatto & G Machado (2009) Efficiency of uniparental male and female care against egg predators in two closely-related syntopic harvestmen. Anim Behav, 78: 1169-1176. [PDF at Anim Behav]

9. Buzatto BA, GS Requena & G Machado (2009) Chemical communication in the gregarious psocid Cerastipsocus sivorii (Psocoptera: Psocidae). J Ins Behav, 22(5): 388-398. [PDF at J Ins Behav]

8. Buzatto BA & G Machado (2009) Amphisexual care in Acutisoma proximum (Arachnida: Opiliones), a neotropical harvestman with exclusive maternal care. Ins Soc, 56(1): 106-108.
[PDF at Ins Soc]

7. Machado G, GS Requena &
BA Buzatto (2009) Reproductive behavior in harvestman (Arachnida): mating systems and parental care. Oecol Bras, 13(1): 58-79.
[PDF (in Portuguese with English and Spanish abstracts)]

2008 and earlier (pre-PhD)

6. Buzatto BA & G Machado (2008) Resource defense polygyny shifts to female defense polygyny over the course of the reproductive season of a neotropical harvestman. Behav Ecol Sociobiol, 63(1): 85-94.
[PDF at Behav Ecol Sociobiol]

5. Contreras-Garduño J,
BA Buzatto, MA Serrano-Meneses, K Nájera-Cordero & A Córdoba-Aguilar (2008) The red wing spot of the American rubyspot (Hetaerina americana) as a heightened condition dependent ornament. Behav Ecol, 19(4): 724-732.
[PDF at Behav Ecol]
Highly cited and featured in a virtual issue of Behav Ecol in honour of Amotz Zahavi [at Behav Ecol]

4. Buzatto BA, GS Requena, EG Martins & G Machado (2007) Effects of maternal care on the lifetime reproductive success of females in a neotropical harvestman. J Anim Ecol, 76(5): 937-945. [PDF at J Anim Ecol]

3. Contreras-Garduño J,
BA Buzatto, L Abundis & A Córdoba-Aguilar (2007) Wing colour properties do not reflect male condition in the American rubyspot (Hetaerina americana). Ethology, 113(10): 944-952.
[PDF at Ethology]

2. Requena GS,
BA Buzatto & G Machado (2007) Habitat use, phenology, and gregariousness of the Neotropical psocopteran Cerastipsocus sivorii (Psocoptera: Psocidae). Sociobiology, 49(3): 197-214.

1. Machado G, GS Requena, BA Buzatto, F Osses & LM Rossetto (2004) Five new cases of paternal care in harvestmen (Arachnida: Opiliones): implications for the evolution of male guarding in the Neotropical family Gonyleptidae. Sociobiology, 44(3): 577-598. [PDF]

Scholarly book chapters

2. Buzatto BA, JL Tomkins & LW Simmons (2014). Alternative phenotypes within mating systems. In The Evolution of Insect Mating Systems. pp. 106-128. (DM Shuker & LW Simmons, Eds), Oxford University Press.

1. Buzatto BA, R. Macías-Ordóñez & G Machado (2013) The macroecology of harvestmen mating systems. In Sexual Selection: Insights from the Neotropics. pp. 115-162. (RHF Macedo & G Machado, Eds), Elsevier.

Other publications

Buzatto BA & RC Firman (2016) Precopulatory Intrasexual Competition. In Encyclopedia of Evolutionary Psychological Science. pp. 1-6. (V Weekes-Shackelford & TK Shackelford, Eds), Springer.